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Lignin

Lignin is a complex organic polymer deposited in the extracellular matrix of plant cell walls, where it fills spaces between cellulose, hemicellulose, and pectin. It confers rigidity and hydrophobicity and helps resist microbial attack.

It is the second most abundant natural polymer on Earth after cellulose. Lignin is highly heterogeneous and

In vascular plants, lignin biosynthesis occurs via the phenylpropanoid pathway in specialized cells such as xylem

Structure features include a range of ether and carbon–carbon linkages, with the β-O-4 linkage being most common.

Function: Lignin provides mechanical strength, reduces cell wall porosity, controls water transport, and acts as a

Industrial relevance: Lignin is a major byproduct of pulping, paper production, and many biorefinery processes. It

Degradation and processing: Lignin is resistant to degradation due to its heterogeneity. Some white-rot fungi and

Variants: Technical lignin refers to lignin isolated from industrial processes; native lignin remains in plant cell

does
not
have
a
single
repeating
unit.
It
is
formed
from
radical
coupling
of
the
monolignols
p-coumaryl
alcohol,
coniferyl
alcohol,
and
sinapyl
alcohol.
tracheary
elements
and
fibers.
The
resulting
polymer
is
deposited
in
the
cell
wall
during
secondary
wall
formation.
The
network
is
highly
cross-linked,
contributing
to
its
recalcitrance
to
chemical
and
enzymatic
breakdown.
barrier
against
pathogens
and
herbivores.
Its
presence
lowers
digestibility
of
woody
biomass
for
ruminants.
is
abundant
and
underutilized,
prompting
research
into
lignin
valorization
to
renewable
chemicals,
fuels,
materials,
or
carbon
fibers.
certain
bacteria
can
depolymerize
lignin.
Pretreatment
methods
help
to
separate
lignin
from
cellulose
and
hemicellulose
for
biofuel
production.
walls.
Analytical
techniques
include
NMR,
thioacidolysis,
and
pyrolysis-GC/MS.