Intine
Intine is the inner layer of the pollen grain wall. It lies inside the outer exine and is in contact with the pollen cytoplasm. The exine is typically thick and resistant due to sporopollenin, while the intine is comparatively thin and more organic in composition. The intine is usually rich in cellulose and pectic substances, and may contain proteins and mucilage. It is formed during pollen development, primarily by material secreted by the pollen grain itself, with additional contributions from surrounding tapetal tissue in some species.
The intine plays a role in pollen hydration and germination. It often forms or thickens near apertures