Plastoquinol
Plastoquinol is a reduced form of plastoquinone, a mobile electron carrier involved in the light-dependent reactions of photosynthesis. It is found in the thylakoid membranes of chloroplasts, where it plays a crucial role in the electron transport chain. Plastoquinol is produced when plastoquinone accepts two electrons and two protons from photosystem I, forming a plastoquinol radical. This process is essential for the generation of the proton gradient that drives ATP synthesis. Plastoquinol then donates its electrons to the cytochrome b6f complex, which further transfers them to plastoquinone, completing the cycle. The oxidation of plastoquinol to plastoquinone is coupled to the reduction of plastoquinone to plastoquinol, facilitating the continuous flow of electrons and protons in the thylakoid membrane. This process is vital for the production of ATP and NADPH, which are used in the Calvin cycle to fix carbon dioxide into organic molecules. Plastoquinol is also involved in the regulation of photosynthesis, as its concentration can influence the activity of the cytochrome b6f complex and the electron transport chain.